Education 1980 B.A. Biochemistry Department, University of California, Berkeley
1987 Ph.D. Biochemistry Department, University of Washington, with Dr. Richard D. Palmiter Thesis: Transmission Ratio Distortion and Mosaicism in Transgenic Mice
Postdoctoral Training 1986-1992 NIH Postdoctoral Fellow with Dr. Melvin I. Simon California Institute of Technology, Pasadena CA
Academic Appointment 1993 Assistant Professor, Department of Pharmacology 2000 Associate Professor, Department of Pharmacology